Title of Paul's PhD Thesis: The Development and Properties of pre-B cells and B-lymphocytes. November 1980.

My PhD Supervisor was Dr HNA Willcox, now of the Institute of Molecular Medicine, University of Oxford. 

He and his wife, Pip, remain fast and valued friends of myself and my wife.

My PhD project involved research on the ontogeny of immunoglobulin producing B-lymphocytes. B-lymphocyte precursors were enriched by density separation methods and found to contain immunoglobulin µ-chains but lack the surface immunoglobulin found in mature B-lymphocytes. My investigations of the surface markers, physical characteristics, responses to mitogens in vitro and location of these cells in the body, as compared with mature B-lymphocytes, formed the basis of a hypothesis of B-lymphocyte development described in publications 3 and 4 in my list of published work..